Multifarious response diversity experiments
Francesco Polazzo, to be added
28 February, 2024
1 Intention
The goal of this document is to calculate the growth rate (r) and the carrying capacity (K) of 6 species of ciliates that we grew individually in a replicated (n = 3) factorial experiment with 5 temperatures (18, 21, 24, 26, and 28), 5 nutrients levels and their combinations (= 25 treatments) for 3 weeks. The calculated r and K are then going to be used as the traits to calculate potential response diversity of communities composed of 2, 3, and 4 species to all possible changes in temperature and nutrients. The calculated potential response diversity will inform us on which communities will be used in the following experimental step. We are going to calculate the growth rate as the slope of the regression of ln(Nt) where: ln is natural log10 and Nt is the population density at time t, during the period of exponential growth. We are going to set initially the period of exponential growth as the first 6 days, but we are going to visually check whether this choice is correct. K will be calculated as the highest population biomass for each population during the experiment.
Let’s start loading the data set and creating a subset for calculating r
## [1] "species" "file" "temperature" "nutrients" "sample_ID"
## [6] "date" "density" "day"## 'data.frame': 119028 obs. of 6 variables:
## $ species : chr "Colpidium" "Colpidium" "Colpidium" "Colpidium" ...
## $ temperature: int 18 18 18 18 18 18 18 18 18 18 ...
## $ nutrients : int 4 2 5 1 4 5 3 2 5 1 ...
## $ sample_ID : chr "C10" "C6" "C15" "C2" ...
## $ density : num 119 164 186 136 282 ...
## $ day : int 1 1 1 1 1 1 1 1 1 1 ...## species temperature nutrients sample_ID density day
## 1 Colpidium 18 4 C10 100 0
## 2 Colpidium 18 4 C10 100 0
## 3 Colpidium 18 4 C10 100 0
## 4 Colpidium 18 4 C10 100 0
## 5 Colpidium 18 4 C10 100 0
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## 16351 Colpidium 18 4 C10 100 0
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## 16376 Colpidium 18 4 C10 100 0
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## 16379 Colpidium 18 4 C10 100 0
## 16380 Colpidium 18 4 C10 100 0
## 16381 Colpidium 18 4 C10 100 0
## 16382 Colpidium 18 4 C10 100 0
## 16383 Colpidium 18 4 C10 100 0
## 16384 Colpidium 18 4 C10 100 0
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## 16386 Colpidium 18 4 C10 100 0
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## 16410 Colpidium 18 4 C10 100 0
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## 16416 Colpidium 18 4 C10 100 0
## 16417 Colpidium 18 4 C10 100 0
## 16418 Colpidium 18 4 C10 100 0
## 16419 Colpidium 18 4 C10 100 0
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## 16421 Colpidium 18 4 C10 100 0
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## 16495 Colpidium 18 4 C10 100 0
## 16496 Colpidium 18 4 C10 100 0
## 16497 Colpidium 18 4 C10 100 0
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## 16500 Colpidium 18 4 C10 100 0
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## 16512 Colpidium 18 4 C10 100 0
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## 16515 Colpidium 18 4 C10 100 0
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## 16517 Colpidium 18 4 C10 100 0
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## 16521 Colpidium 18 4 C10 100 0
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## 16523 Colpidium 18 4 C10 100 0
## 16524 Colpidium 18 4 C10 100 0
## 16525 Colpidium 18 4 C10 100 0
## 16526 Colpidium 18 4 C10 100 0
## 16527 Colpidium 18 4 C10 100 0
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## 16530 Colpidium 18 4 C10 100 0
## 16531 Colpidium 18 4 C10 100 0
## 16532 Colpidium 18 4 C10 100 0
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## 16544 Colpidium 18 4 C10 100 0
## 16545 Colpidium 18 4 C10 100 0
## 16546 Colpidium 18 4 C10 100 0
## 16547 Colpidium 18 4 C10 100 0
## 16548 Colpidium 18 4 C10 100 0
## 16549 Colpidium 18 4 C10 100 0
## 16550 Colpidium 18 4 C10 100 0
## 16551 Colpidium 18 4 C10 100 0
## 16552 Colpidium 18 4 C10 100 0
## 16553 Colpidium 18 4 C10 100 0
## 16554 Colpidium 18 4 C10 100 0
## 16555 Colpidium 18 4 C10 100 0
## 16556 Colpidium 18 4 C10 100 0
## 16557 Colpidium 18 4 C10 100 0
## 16558 Colpidium 18 4 C10 100 0
## 16559 Colpidium 18 4 C10 100 0
## 16560 Colpidium 18 4 C10 100 0
## 16561 Colpidium 18 4 C10 100 0
## 16562 Colpidium 18 4 C10 100 0
## 16563 Colpidium 18 4 C10 100 0
## 16564 Colpidium 18 4 C10 100 0
## 16565 Colpidium 18 4 C10 100 0
## 16566 Colpidium 18 4 C10 100 0
## 16567 Colpidium 18 4 C10 100 0
## 16568 Colpidium 18 4 C10 100 0
## 16569 Colpidium 18 4 C10 100 0
## 16570 Colpidium 18 4 C10 100 0
## 16571 Colpidium 18 4 C10 100 0
## 16572 Colpidium 18 4 C10 100 0
## 16573 Colpidium 18 4 C10 100 0
## 16574 Colpidium 18 4 C10 100 0
## 16575 Colpidium 18 4 C10 100 0
## 16576 Colpidium 18 4 C10 100 0
## 16577 Colpidium 18 4 C10 100 0
## 16578 Colpidium 18 4 C10 100 0
## 16579 Colpidium 18 4 C10 100 0
## 16580 Colpidium 18 4 C10 100 0
## 16581 Colpidium 18 4 C10 100 0
## 16582 Colpidium 18 4 C10 100 0
## 16583 Colpidium 18 4 C10 100 0
## 16584 Colpidium 18 4 C10 100 0
## 16585 Colpidium 18 4 C10 100 0
## 16586 Colpidium 18 4 C10 100 0
## 16587 Colpidium 18 4 C10 100 0
## 16588 Colpidium 18 4 C10 100 0
## 16589 Colpidium 18 4 C10 100 0
## 16590 Colpidium 18 4 C10 100 0
## 16591 Colpidium 18 4 C10 100 0
## 16592 Colpidium 18 4 C10 100 0
## 16593 Colpidium 18 4 C10 100 0
## 16594 Colpidium 18 4 C10 100 0
## 16595 Colpidium 18 4 C10 100 0
## 16596 Colpidium 18 4 C10 100 0
## 16597 Colpidium 18 4 C10 100 0
## 16598 Colpidium 18 4 C10 100 0
## 16599 Colpidium 18 4 C10 100 0
## 16600 Colpidium 18 4 C10 100 0
## 16601 Colpidium 18 4 C10 100 0
## 16602 Colpidium 18 4 C10 100 0
## 16603 Colpidium 18 4 C10 100 0
## 16604 Colpidium 18 4 C10 100 0
## 16605 Colpidium 18 4 C10 100 0
## 16606 Colpidium 18 4 C10 100 0
## 16607 Colpidium 18 4 C10 100 0
## 16608 Colpidium 18 4 C10 100 0
## 16609 Colpidium 18 4 C10 100 0
## 16610 Colpidium 18 4 C10 100 0
## 16611 Colpidium 18 4 C10 100 0
## 16612 Colpidium 18 4 C10 100 0
## 16613 Colpidium 18 4 C10 100 0
## 16614 Colpidium 18 4 C10 100 0
## 16615 Colpidium 18 4 C10 100 0
## 16616 Colpidium 18 4 C10 100 0
## 16617 Colpidium 18 4 C10 100 0
## 16618 Colpidium 18 4 C10 100 0
## 16619 Colpidium 18 4 C10 100 0
## 16620 Colpidium 18 4 C10 100 0
## 16621 Colpidium 18 4 C10 100 0
## 16622 Colpidium 18 4 C10 100 0
## 16623 Colpidium 18 4 C10 100 0
## 16624 Colpidium 18 4 C10 100 0
## 16625 Colpidium 18 4 C10 100 0
## 16626 Colpidium 18 4 C10 100 0
## 16627 Colpidium 18 4 C10 100 0
## 16628 Colpidium 18 4 C10 100 0
## 16629 Colpidium 18 4 C10 100 0
## 16630 Colpidium 18 4 C10 100 0
## 16631 Colpidium 18 4 C10 100 0
## 16632 Colpidium 18 4 C10 100 0
## 16633 Colpidium 18 4 C10 100 0
## 16634 Colpidium 18 4 C10 100 0
## 16635 Colpidium 18 4 C10 100 0
## 16636 Colpidium 18 4 C10 100 0
## 16637 Colpidium 18 4 C10 100 0
## 16638 Colpidium 18 4 C10 100 0
## 16639 Colpidium 18 4 C10 100 0
## 16640 Colpidium 18 4 C10 100 0
## 16641 Colpidium 18 4 C10 100 0
## 16642 Colpidium 18 4 C10 100 0
## 16643 Colpidium 18 4 C10 100 0
## 16644 Colpidium 18 4 C10 100 0
## 16645 Colpidium 18 4 C10 100 0
## 16646 Colpidium 18 4 C10 100 0
## 16647 Colpidium 18 4 C10 100 0
## 16648 Colpidium 18 4 C10 100 0
## 16649 Colpidium 18 4 C10 100 0
## 16650 Colpidium 18 4 C10 100 0
## 16651 Colpidium 18 4 C10 100 0
## 16652 Colpidium 18 4 C10 100 0
## 16653 Colpidium 18 4 C10 100 0
## 16654 Colpidium 18 4 C10 100 0
## 16655 Colpidium 18 4 C10 100 0
## 16656 Colpidium 18 4 C10 100 0
## 16657 Colpidium 18 4 C10 100 0
## 16658 Colpidium 18 4 C10 100 0
## 16659 Colpidium 18 4 C10 100 0
## 16660 Colpidium 18 4 C10 100 0
## 16661 Colpidium 18 4 C10 100 0
## 16662 Colpidium 18 4 C10 100 0
## 16663 Colpidium 18 4 C10 100 0
## 16664 Colpidium 18 4 C10 100 0
## 16665 Colpidium 18 4 C10 100 0
## 16666 Colpidium 18 4 C10 100 0
## [ reached 'max' / getOption("max.print") -- omitted 697502 rows ]
(#fig:time_series_spp)Time series of species densities across the treatments.
m, , h 00## Calculate K and r Visual inspection of exponential growth phase
(#fig:reg_all)Species densities across the treatments with the regression lines used to calculate the intrinsic rate of growth (r).
1.1 GAMs to fit responde surface
At least for now, we focus on the intrinsic rate of growth (r). We are going to fit response surfaces using the calculated r for all species using GAMs. Then we are also going to use r as the species’ trait to calculate potential response diversity, as well as response diversity when the direction of the environmental change is known. The rationale is that r is likely of more relevance if the environmental change of interest occurs rapidly, since r provides information on a population’s ability to rapidly bounce back after disturbance. In the upcoming experiment, we will have temperature fluctuating relatively fast, and so we decide now to focus on r.
Use GAMs to fit response surface of r and K
species temperature nutrients predicted1 Colpidium 18 1.00 0.06013854 2 Colpidium 18 1.01 0.06160047 3 Colpidium 18 1.02 0.06306235 4 Colpidium 18 1.03 0.06452412 5 Colpidium 18 1.04 0.06598574 6 Colpidium 18 1.05 0.06744714
Create surface plots
We now check how the GAMs surfaces (r) look compared to the measured densities.
Checking predictions
(#fig:surface_Dexi)Measured density values of Dexiostoma in the different treatments (a) vs fitted surface of growth rate (b).
(#fig:surface_colp)Measured density values of Colpidium in the different treatments (a) vs fitted surface of growth rate (b).
(#fig:surface_loxo)Measured density values of Loxocephalus in the different treatments (a) vs fitted surface of growth rate (b).
(#fig:surface_paramecium)Measured density values of Paramecium in the different treatments (a) vs fitted surface of growth rate (b).
(#fig:surface_spiro)Measured density values of Spirostotum in the different treatments (a) vs fitted surface of growth rate (b).
(#fig:surface_tetra)Measured density values of Tetrahymena in the different treatments (a) vs fitted surface of growth rate (b).
2 Calculate Potential Response diversity
After discussion, we decided to remove Tetrahymena
Now, we calculate potential response diversity for all possible communities composed of 2, 3, and 4 species. We start with all communities composed of 2 species.
2.1 Potential response diversity 2 species communities
Steps are:
Create ref environmental conditions
Create all possible communities with 2 species
Calculate potential response diversity for all communities richness = 2 with our function “get_potential_RD”
Results divergence dissimilarity species composition richness 1 0.20489479 1.044523 Colpidium_Dexiostoma CD 2 2 0.13437387 1.035102 Colpidium_Loxocephalus CL 2 3 0.15169106 1.039528 Colpidium_Paramecium CP 2 4 0.03583113 1.028196 Colpidium_Spirostomum CS 2 5 0.14136269 1.037479 Dexiostoma_Loxocephalus DL 2 6 0.24765575 1.060830 Dexiostoma_Paramecium DP 2
2.2 Potential Response Diversity 3 species communities
Results divergence dissimilarity species composition 1 0.2580088 1.052634 Colpidium_Dexiostoma_Loxocephalus CDL 2 0.3036517 1.065257 Colpidium_Dexiostoma_Paramecium CDP 3 0.2285406 1.054633 Colpidium_Dexiostoma_Spirostomum CDS 4 0.2155091 1.054397 Colpidium_Loxocephalus_Paramecium CLP 5 0.1485689 1.045846 Colpidium_Loxocephalus_Spirostomum CLS 6 0.1662250 1.049642 Colpidium_Paramecium_Spirostomum CPS richness 1 3 2 3 3 3 4 3 5 3 6 3
2.3 Potential Response Diversity 4 species communities
Results
divergence dissimilarity species 1 0.3351715 1.067317 Colpidium_Dexiostoma_Loxocephalus_Paramecium 2 0.2693354 1.059176 Colpidium_Dexiostoma_Loxocephalus_Spirostomum 3 0.3168594 1.067527 Colpidium_Dexiostoma_Paramecium_Spirostomum 4 0.2237600 1.058784 Colpidium_Loxocephalus_Paramecium_Spirostomum 5 0.3313425 1.070155 Dexiostoma_Loxocephalus_Paramecium_Spirostomum composition richness 1 CDLP 4 2 CDLS 4 3 CDPS 4 4 CLPS 4 5 DLPS 4
(#fig:potential_RD_plot)Potential Response diversity per species richness measured as dissimilarity (a) and divergence (b).
3 Response diversity with known direction of environmental change
Now we calculate RD with known direction of environmental change, i.e. the ones we are going to apply in the experiment. We are going to calculate response diversity for multiple possible ranges of temperature fluctuations. - Create a data set with environmental conditions that we may use in the experiment and calculate RD knowing the direction of the environmental change.
- fluctuating temperature x 3 fixed nutrients = 9 treatments
3.1 Magnitude of T change - 1
- first fluctuation ranges are going to be 18-22, 22-26, and 24 - 28. Temperature stays at the lower end of the range (i.e. 18C) for 3 days, it increases over 24h and stays at the higher end for 3 days.
(#fig:env_sp2_t1)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.1.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation (since temperature and nutrients are the same for 3 days in a row, the magnitude of the unit vector is zero [no direction of environmental change], so we get NAs for divergence. We replace these with zeros)
Results time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_22_Nut1 18 1 0 NaN 2 1 Mag_18_22_Nut3 18 3 0 NaN 3 1 Mag_18_22_Nut5 18 5 0 NaN 4 1 Mag_22_26_Nut1 22 1 0 NaN 5 1 Mag_22_26_Nut3 22 3 0 NaN 6 1 Mag_22_26_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2 [1] “CD” “CL” “CP” “CS” “DL” “DP” “DS” “LP” “LS” “PS”
(#fig:RD_2sp_t1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.1.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_22_Nut1 18 1 0 NaN 2 1 Mag_18_22_Nut3 18 3 0 NaN 3 1 Mag_18_22_Nut5 18 5 0 NaN 4 1 Mag_22_26_Nut1 22 1 0 NaN 5 1 Mag_22_26_Nut3 22 3 0 NaN 6 1 Mag_22_26_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_t1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.1.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_22_Nut1 18 1 0 NaN 2 1 Mag_18_22_Nut3 18 3 0 NaN 3 1 Mag_18_22_Nut5 18 5 0 NaN 4 1 Mag_22_26_Nut1 22 1 0 NaN 5 1 Mag_22_26_Nut3 22 3 0 NaN 6 1 Mag_22_26_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_t1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_t1)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.2 Magnitude of T change - 2
- 3C magnitude
- fluctuation ranges are going to be 18-21, 22-25, and 25 - 28
(#fig:env_scenario2_t2)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.2.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_21_Nut1 18 1 0 NaN 2 1 Mag_18_21_Nut3 18 3 0 NaN 3 1 Mag_18_21_Nut5 18 5 0 NaN 4 1 Mag_22_25_Nut1 22 1 0 NaN 5 1 Mag_22_25_Nut3 22 3 0 NaN 6 1 Mag_22_25_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_t2)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.2.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_21_Nut1 18 1 0 NaN 2 1 Mag_18_21_Nut3 18 3 0 NaN 3 1 Mag_18_21_Nut5 18 5 0 NaN 4 1 Mag_22_25_Nut1 22 1 0 NaN 5 1 Mag_22_25_Nut3 22 3 0 NaN 6 1 Mag_22_25_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_t2)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.2.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_21_Nut1 18 1 0 NaN 2 1 Mag_18_21_Nut3 18 3 0 NaN 3 1 Mag_18_21_Nut5 18 5 0 NaN 4 1 Mag_22_25_Nut1 22 1 0 NaN 5 1 Mag_22_25_Nut3 22 3 0 NaN 6 1 Mag_22_25_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_t2)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_t2)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.3 Magnitude of T change - 3
- 2C magnitude
- third fluctuation ranges are going to be 18-20, 22-24, and 26 - 28
(#fig:env_scenario2_t3)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.3.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_20_Nut1 18 1 0 NaN 2 1 Mag_18_20_Nut3 18 3 0 NaN 3 1 Mag_18_20_Nut5 18 5 0 NaN 4 1 Mag_22_24_Nut1 22 1 0 NaN 5 1 Mag_22_24_Nut3 22 3 0 NaN 6 1 Mag_22_24_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_t3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.3.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_20_Nut1 18 1 0 NaN 2 1 Mag_18_20_Nut3 18 3 0 NaN 3 1 Mag_18_20_Nut5 18 5 0 NaN 4 1 Mag_22_24_Nut1 22 1 0 NaN 5 1 Mag_22_24_Nut3 22 3 0 NaN 6 1 Mag_22_24_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_t3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.3.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Mag_18_20_Nut1 18 1 0 NaN 2 1 Mag_18_20_Nut3 18 3 0 NaN 3 1 Mag_18_20_Nut5 18 5 0 NaN 4 1 Mag_22_24_Nut1 22 1 0 NaN 5 1 Mag_22_24_Nut3 22 3 0 NaN 6 1 Mag_22_24_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_t3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_t3)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.4 Fluctuation frequency - 1
Same temperature fluctuations as before, but now fluctuating over 5 days
Create a data set with environmental conditions that we may use in the experiment and calculate RD knowing the direction of the environmental change.
fluctuating temperature x 3 fixed nutrients = 9 treatments
first fluctuation ranges are going to be 18-22, 22-26, and 24 - 28
(#fig:env_sp2_T1)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.4.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Fluct_18_22_Nut1 18 1 0 NaN 2 1 Fluct_18_22_Nut3 18 3 0 NaN 3 1 Fluct_18_22_Nut5 18 5 0 NaN 4 1 Fluct_22_26_Nut1 22 1 0 NaN 5 1 Fluct_22_26_Nut3 22 3 0 NaN 6 1 Fluct_22_26_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_T1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.4.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Fluct_18_22_Nut1 18 1 0 NaN 2 1 Fluct_18_22_Nut3 18 3 0 NaN 3 1 Fluct_18_22_Nut5 18 5 0 NaN 4 1 Fluct_22_26_Nut1 22 1 0 NaN 5 1 Fluct_22_26_Nut3 22 3 0 NaN 6 1 Fluct_22_26_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_T1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.4.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment E1_ref E2_ref dissimilarity divergence 1 1 Fluct_18_22_Nut1 18 1 0 NaN 2 1 Fluct_18_22_Nut3 18 3 0 NaN 3 1 Fluct_18_22_Nut5 18 5 0 NaN 4 1 Fluct_22_26_Nut1 22 1 0 NaN 5 1 Fluct_22_26_Nut3 22 3 0 NaN 6 1 Fluct_22_26_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_T1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_T1)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.5 Fluctuation frequency - 2
- first fluctuation ranges are going to be 18-21, 22-25, and 25 - 28
(#fig:env_scenario2_T2)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.5.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Fluct_18_21_Nut1 18 1 0 NaN 2 1 Fluct_18_21_Nut3 18 3 0 NaN 3 1 Fluct_18_21_Nut5 18 5 0 NaN 4 1 Fluct_22_25_Nut1 22 1 0 NaN 5 1 Fluct_22_25_Nut3 22 3 0 NaN 6 1 Fluct_22_25_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_T2)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.5.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Fluct_18_21_Nut1 18 1 0 NaN 2 1 Fluct_18_21_Nut3 18 3 0 NaN 3 1 Fluct_18_21_Nut5 18 5 0 NaN 4 1 Fluct_22_25_Nut1 22 1 0 NaN 5 1 Fluct_22_25_Nut3 22 3 0 NaN 6 1 Fluct_22_25_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_T2)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.5.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment E1_ref E2_ref dissimilarity divergence 1 1 Fluct_18_21_Nut1 18 1 0 NaN 2 1 Fluct_18_21_Nut3 18 3 0 NaN 3 1 Fluct_18_21_Nut5 18 5 0 NaN 4 1 Fluct_22_25_Nut1 22 1 0 NaN 5 1 Fluct_22_25_Nut3 22 3 0 NaN 6 1 Fluct_22_25_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_T2)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_T2)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.6 Fluctuation frequency - 3
- third fluctuation ranges are going to be 18-20, 21-24, and 26 - 28
(#fig:env_scenario2_T3)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.6.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Fluct_18_20_Nut1 18 1 0 NaN 2 1 Fluct_18_20_Nut3 18 3 0 NaN 3 1 Fluct_18_20_Nut5 18 5 0 NaN 4 1 Fluct_22_24_Nut1 22 1 0 NaN 5 1 Fluct_22_24_Nut3 22 3 0 NaN 6 1 Fluct_22_24_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_T3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.6.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment E1_ref E2_ref dissimilarity divergence 1 1 Fluct_18_20_Nut1 18 1 0 NaN 2 1 Fluct_18_20_Nut3 18 3 0 NaN 3 1 Fluct_18_20_Nut5 18 5 0 NaN 4 1 Fluct_22_24_Nut1 22 1 0 NaN 5 1 Fluct_22_24_Nut3 22 3 0 NaN 6 1 Fluct_22_24_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_T3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.6.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Fluct_18_20_Nut1 18 1 0 NaN 2 1 Fluct_18_20_Nut3 18 3 0 NaN 3 1 Fluct_18_20_Nut5 18 5 0 NaN 4 1 Fluct_22_24_Nut1 22 1 0 NaN 5 1 Fluct_22_24_Nut3 22 3 0 NaN 6 1 Fluct_22_24_Nut5 22 5 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_T3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_T3)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.7 Changing nutrients - 1
Same temperature fluctuations as before, different nutrient levels
Create a data set with environmental conditions that we may use in the experiment and calculate RD knowing the direction of the environmental change.
fluctuating temperature x 3 fixed nutrients = 9 treatments
first fluctuation ranges are going to be 18-22, 22-26, and 24 - 28
(#fig:env_sp2_nut)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.7.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_22_Nut1 18 1 0 NaN 2 1 Nut_18_22_Nut2 18 2 0 NaN 3 1 Nut_18_22_Nut4 18 4 0 NaN 4 1 Nut_22_26_Nut1 22 1 0 NaN 5 1 Nut_22_26_Nut2 22 2 0 NaN 6 1 Nut_22_26_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_nut1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.7.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_22_Nut1 18 1 0 NaN 2 1 Nut_18_22_Nut2 18 2 0 NaN 3 1 Nut_18_22_Nut4 18 4 0 NaN 4 1 Nut_22_26_Nut1 22 1 0 NaN 5 1 Nut_22_26_Nut2 22 2 0 NaN 6 1 Nut_22_26_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_nut1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.7.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_22_Nut1 18 1 0 NaN 2 1 Nut_18_22_Nut2 18 2 0 NaN 3 1 Nut_18_22_Nut4 18 4 0 NaN 4 1 Nut_22_26_Nut1 22 1 0 NaN 5 1 Nut_22_26_Nut2 22 2 0 NaN 6 1 Nut_22_26_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_nut1)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_nut1)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.8 Changing nutrients - 2
- first fluctuation ranges are going to be 18-21, 22-25, and 25 - 28
(#fig:env_scenario2_nut)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.8.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_21_Nut1 18 1 0 NaN 2 1 Nut_18_21_Nut2 18 2 0 NaN 3 1 Nut_18_21_Nut4 18 4 0 NaN 4 1 Nut_22_25_Nut1 22 1 0 NaN 5 1 Nut_22_25_Nut2 22 2 0 NaN 6 1 Nut_22_25_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_nut)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.8.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_21_Nut1 18 1 0 NaN 2 1 Nut_18_21_Nut2 18 2 0 NaN 3 1 Nut_18_21_Nut4 18 4 0 NaN 4 1 Nut_22_25_Nut1 22 1 0 NaN 5 1 Nut_22_25_Nut2 22 2 0 NaN 6 1 Nut_22_25_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_nut)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.8.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_21_Nut1 18 1 0 NaN 2 1 Nut_18_21_Nut2 18 2 0 NaN 3 1 Nut_18_21_Nut4 18 4 0 NaN 4 1 Nut_22_25_Nut1 22 1 0 NaN 5 1 Nut_22_25_Nut2 22 2 0 NaN 6 1 Nut_22_25_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_nut)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_nut)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
3.9 Changing nutrients - 3
- third fluctuation ranges are going to be 18-20, 22-24, and 26 - 28
(#fig:env_scenario3_nut)Time series of the environmental conditions for temperature (a) and nutrients (b). These environmental conditions represent the “treatments” and will be crossed in a full factorial design resulting in 9 unique treatments
3.9.1 Response diversity 2 species communities
Same steps as before:
Create all possible communities with 2 species
Actual RD calculation
Results time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_20_Nut1 18 1 0 NaN 2 1 Nut_18_20_Nut2 18 2 0 NaN 3 1 Nut_18_20_Nut4 18 4 0 NaN 4 1 Nut_22_24_Nut1 22 1 0 NaN 5 1 Nut_22_24_Nut2 22 2 0 NaN 6 1 Nut_22_24_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma CD 2 2 Colpidium_Dexiostoma CD 2 3 Colpidium_Dexiostoma CD 2 4 Colpidium_Dexiostoma CD 2 5 Colpidium_Dexiostoma CD 2 6 Colpidium_Dexiostoma CD 2
(#fig:RD_2sp_nut3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 2 species
3.9.2 Response diversity 3 species communities
Now, we calculate RD for all possible communities of 3 species knowing the direction of the environmental change (we created before).
time treatment E1_ref E2_ref dissimilarity divergence 1 1 Nut_18_20_Nut1 18 1 0 NaN 2 1 Nut_18_20_Nut2 18 2 0 NaN 3 1 Nut_18_20_Nut4 18 4 0 NaN 4 1 Nut_22_24_Nut1 22 1 0 NaN 5 1 Nut_22_24_Nut2 22 2 0 NaN 6 1 Nut_22_24_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus CDL 3 2 Colpidium_Dexiostoma_Loxocephalus CDL 3 3 Colpidium_Dexiostoma_Loxocephalus CDL 3 4 Colpidium_Dexiostoma_Loxocephalus CDL 3 5 Colpidium_Dexiostoma_Loxocephalus CDL 3 6 Colpidium_Dexiostoma_Loxocephalus CDL 3
(#fig:RD_3sp_nut3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 3 species
3.9.3 Response diversity 4 species communities
Now, we calculate RD for all possible communities of 4 species knowing the direction of the environmental change (we created before).
time treatment temperature nutrients dissimilarity divergence 1 1 Nut_18_20_Nut1 18 1 0 NaN 2 1 Nut_18_20_Nut2 18 2 0 NaN 3 1 Nut_18_20_Nut4 18 4 0 NaN 4 1 Nut_22_24_Nut1 22 1 0 NaN 5 1 Nut_22_24_Nut2 22 2 0 NaN 6 1 Nut_22_24_Nut4 22 4 0 NaN species composition richness 1 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 2 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 3 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 4 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 5 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4 6 Colpidium_Dexiostoma_Loxocephalus_Paramecium CDLP 4
(#fig:RD_4sp_nut3)Response diversity calcualted as dissimilarity (a) and diversity (b) of communities composed of all possible combinations of 4 species
(#fig:summary_RD_nut3)Summary of Response divergence measured for all possible communities of 2, 3, and 4 species with known direction of environmental change
4 Big Results table
## [1] "time" "treatment" "temperature" "nutrients"
## [5] "dissimilarity" "divergence" "species" "composition"
## [9] "richness"## [1] "divergence" "dissimilarity" "species" "composition"
## [5] "richness"